Departments & Positions
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Command
The Command department is ultimately responsible for the ship and its crew, and those within the department are responsible for commanding the vessel and representing the interests of Starfleet.
Commanding Officer Ultimately responsible for the ship and crew, the Commanding Officer is the most senior officer aboard a vessel. S/he is responsible for carrying out the orders of Starfleet, and for representing both Starfleet and the Federation.
When playing the Commanding Officer think to yourself about Kirk's role.

First Officer The liaison between captain and crew, the First Officer acts as the disciplinarian, personnel manager, advisor to the captain, and much more. S/he is also one of only two officers, along with the Chief Medical Officer, that can remove a Commanding Officer from duty.
Only exceptional applications for this position will be accepted.
When playing the First Officer think to yourself about Spock's role.

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Operations
Responsible for the navigation and flight control of a vessel and its auxiliary craft, the Operations division includes pilots trained in both starship and auxiliary craft piloting as well as navigators and other personnel to help in running the various ancillary operations of the ship.
Chief Helmsman/Tactical Officer A Helm Officer must always be present on the bridge of a starship. S/he plots courses, supervises the computers piloting, corrects any flight deviations and pilots the ship manually when needed. The Chief Helmsman is the senior most Helm Officer aboard, serving as a Senior Officer, chief of the personnel under him/her, and handles all weapons aboard the ship.
When playing the Chief Helmsman and Tactical Officer think to yourself about Sulu's role.

Chief Navigator/Security Officer The Chief Navigator is the senior most navigator on the ship and is responsible for all navigators aboard the vessel. Responsibilities of the navigators is include projecting the course of a starship and determining a ship's position, velocity and direction in relationship to a course. The navigator can also use the ship's navigational sensors to determine the positions, speeds and trajectories of other objects. Additionally, this officer is in charge of the security of the ship and crew.
When playing the Chief Navigator and Security Officer think to yourself about Chekov's role.

Chief Communications Officer Responsible for maintaining and upgrading the universal translator, controls the intercom and responsible for coordinating communications with other ships, stations or colonies/planets.
When playing the Chief Communications Officer think to yourself about Uhura's role.

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Engineering
The engineering division has the enormous task of keeping the ship working; they are responsible for making repairs, fixing problems, and making sure that the ship is ready for anything.
Chief Engineer The Chief Engineer is responsible for the condition of all systems and equipment on board a Starfleet ship or facility. S/he oversees maintenance, repairs and upgrades of all equipment. S/he is also responsible for the many repairs teams during crisis situations.
The Chief Engineer is not only the department head but also a senior officer, responsible for all the crew members in her/his department and maintenance of the duty rosters.
When playing the Chief Engineering Officer think to yourself about Scotty's role.

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Science
From sensor readings to figuring out a way to enter the strange spacial anomaly, the science division is responsible for recording data, testing new ideas out, and making discoveries.
Chief Science Officer The Chief Science Officer is responsible for all the scientific data the ship/facility collects, and the distribution of such data to specific section within the department for analysis. S/he is also responsible with providing the ship's captain with scientific information needed for command decisions.
S/he also is a department head and a member of the Senior Staff and responsible for all the crew members in her/his department and duty rosters.
When playing the Chief Science Officer think to yourself about Spock's role.

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Medical
The medical division is responsible for the mental and physical health of the crew, from running annual physicals to combatting a strange plague that is afflicting the crew to helping a crew member deal with the loss of a loved one.
Chief Medical Officer The Chief Medical Officer is responsible for the physical health of the entire crew, but does more than patch up injured crew members. His/her function is to ensure that they do not get sick or injured to begin with, and to this end monitors their health and conditioning with regular check ups. If necessary, the Chief Medical Officer can remove anyone from duty, even a Commanding Officer. Besides this s/he is available to provide medical advice to any individual who requests it.
When playing the Chief Medical Officer think to yourself about McCoy's role.
